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About signal flies

Lophoplatystoma molle Hendel, 1914

Diagnostic characters of Platystomatidae

Platystomatidae are considered monophyletic from a morphological (McAlpine 1989 and Korneyev 1999) and molecular (Han and Ro 2005) perspective, but the lineage is weakly supported by the Minimum Evolution tree, and there are indications that Platystomatidae may be a paraphyletic group basal to the Ulidiidae.

The diagnostic characters (synapomorphies) common to all Platystomatidae, that separate them from these families are listed below (the most significant in bold typeface).

Synapomorphies of Platystomatidae

  • ocelli present (except Bromophila)
  • fronto-orbital setae absent
  • genal setae present, but true vibrissae are absent
  • R1 & R4+5 with dorsal setae
  • Wing cell cua  closed by a straight vein,  perpendicular to vein CuA
  • internal sternal apodemes of sternites 4-6 are absent.

Platystomatidae in unsorted material

Elassogaster Bigot, 1860

Sorting Platystomatidae

 

Adult Platystomatidae are frequently confused with other higher Tephritoidea: Tephritidae (or fruit flies), the Ulidiidae (picture-winged flies), the Pyrgotidae (waved light flies) and the Richardiidae. They can be sorted from these families using the key characters described below.


key characters: platystomatidae

Absence of fronto-orbital setae

Square posterior apical angle of anterior cubital cell (cua)

Square posterior apical angle of anterior cubital cell (cua)

Differentiating Platystomatidae from Ulidiidae: features of the head.

Fronto-orbital setae (fr s) are located along  the upper orbital margin; genal setae (gn s) and ocelli (oc) are also shown.

Square posterior apical angle of anterior cubital cell (cua)

Square posterior apical angle of anterior cubital cell (cua)

Square posterior apical angle of anterior cubital cell (cua)

Differentiating Platystomatidae from Ulidiidae: anal cell of the wing

Lack of any extension of the posterior apical angle of anterior cubital cell (cua)

Presence of setae along certain wing veins

Square posterior apical angle of anterior cubital cell (cua)

Presence of setae along certain wing veins

Differentiating Platystomatidae: veins of the wing

The presence of setae along the entire length of wing vein R1, frequently vein R4+5 and sometimes the basal portion of vein M4, where it is adjacent to cells bm and cua


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